All trees
Sword
Greatsword
Blunt
Bow
Fire Magic
Water Magic
Nature Magic
Shield
Spear
Warrior
Guardian
Fortress
Burst
Blitz
Execution
Berserker
Tide
Harmony
Stake
SwordGreatswordBluntBowFire MagicWater MagicNature MagicShieldSpearWarriorGuardianFortressBurstBlitzExecutionBerserkerTideHarmonyStakeHarmony Skill Tree overview
Enhances the supporting ability of nature magic.
From backgrounds (7): 
Mage, 
Forest Keeper, 
Hunter, 
Gardener, 
Druid, 
Prodigy, 
Cursed Child
basic weapon attack (mandatory root) · active · passive P - lines show unlock requirements · top-right corner = energy cost per cast (skill-point cost shows on hover)
All Harmony skills, in full (5)
Inner Grove
SKILL_InnerGrove · Passive Physical
All health recovery effects you receive are increased by 25%.
PositionTier 1, column 1Skill points5
Numbers
| Range | 1 tile |
| Cooldown | 0s |
| Energy cost | 0 |
Life Burst
SKILL_LifeBurst · Passive Physical
When the Strange Seed is destroyed, it explodes, restoring Current Health equal to 200% of Magic Attack Power to all adjacent allied units and granting Cooldown Reduction for 10 seconds.
PositionTier 1, column 2Skill points5
Effects & keywords
Cooldown Reduction (on allies): +30 Cooldown Reduction
Numbers
| Range | 1 tile |
| Cooldown | 0s |
| Energy cost | 0 |
Damage
Shown for stock Perion, Nature Wooden Staff at level 5 (level cap). Pick your physical and magic units in the sidebar.
Heal Restores 196 health to an ally per cast (about 125.7/s at this cadence)
Numbers are before target resistance (damage x 100 / (100 + R); most monsters have 0, armored ones up to 60 for a 37.5% cut) and before hit chance, dodge and guard. Combat difficulty does not change your own damage.
Healing Touch
SKILL_HealingTouch · Passive Physical
When equipped with a nature staff, the basic attack changes to Healing Touch.
PositionTier 1, column 3Skill points5
Effects & keywords
Healing Touch: Restores an ally unit's Current Health by 50% of Magic Attack Power.
Numbers
| Range | 1 tile |
| Cooldown | 0s |
| Energy cost | 0 |
Life Cycle
SKILL_LifeCycle · Passive Physical
When an enemy unit dies during combat, grants Health Boost equal to 1% of Magic Attack Power to all friendly units for 2 seconds.
Effects & keywords
Health Boost (on units in the area): Restores 10 Current Health per stack every second.
Numbers
| Range | 1 tile |
| Cooldown | 0s |
| Energy cost | 0 |
Damage
Shown for stock Perion, Nature Wooden Staff at level 5 (level cap). Pick your physical and magic units in the sidebar.
Expected per cast 1 magic per target
= MAtk 98.2 x 1% x 1.079 crit
Versus benchmark 
0.2% of Acid Slogel HP
0.2% of Acid Slogel HPDamage per second 0.7 per second (at attack speed)
Versus median 0x the median magic active per cast
Numbers are before target resistance (damage x 100 / (100 + R); most monsters have 0, armored ones up to 60 for a 37.5% cut) and before hit chance, dodge and guard. Combat difficulty does not change your own damage.
Green Shade
SKILL_GreenShade · Passive Physical
Damage Taken decreases by 8% for each allied plant structure within 1 tile. (Applies up to a maximum of 40%)
Numbers
| Range | 1 tile |
| Cooldown | 0s |
| Energy cost | 0 |